Hi,
1) Latest from Symantec means the definitions avaialable on the internet.
2) Latest on Manager means the definitions which are currently loaded into the manager.
First one needs internet access so that it can go to web and look for latest one.
Attached screenshot is too small to see the picture. Anyways as long as Latest from Symantec and Latest on Manager is on the same date there shouldn't be anything to worry about.